New Roman’,’serif’; mso-fareast-font-family: 宋体; mso-font-kerning: 0pt">Exceptionally well preserved vegetative shoots of the extinct cheirolepidiaceous conifer Pseudo-frenelopsis parceramosa (Fontaine) Watson are described in this paper from Hauterivian strata within the marine Early Cretaceous Speeton Clay Formation of East Yorkshire in the northern England. As a result of extensive pyritisation the shoots are preserved with little or no compaction in three dimensions and much of the internal anatomy is preserved-particularly of the woody (xylem) tissues. Externally the twigs are enclosed by their original-non-pyritised-cuticle, therefore also preserving anatomical details but in this case mainly of the outer and lateral walls of the epidermis, including stomatal features, and of the hypodermis. This study extends and complements previous studies on P. parceramosa from the largely terrestrial beds of southern England and from elsewhere in the Lower Cretaceous, showing that the Speeton material is a local variant and may possibly represent a different biological species.