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The ionization properties of cardiolipin and its variants in model bilayers
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Cardiolipin ionization was analyzed by independent biophysical approaches.

Both cardiolipin headgroup phosphates behave as strong acids with low pKa values.

Cardiolipin exists as a dianion in the physiological pH range.

20">Ionization is not altered in variants lacking an acyl chain or the central hydroxyl.

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